Eupatorium
Eupatorium is a genus of flowering plants, containing from 36 to 60 species depending on the classification system. Most are herbaceous perennial plants growing to 0.5–3 m tall. A few are shrubs. The genus is native to temperate regions of the Northern Hemisphere. Most are commonly called bonesets, thoroughworts or "snakeroots".
Systematics and taxonomy
Eupatorium has at times been held to contain as many as 800 species[1], but many of these have been moved (at least by some authors) to other genera, including Ageratina, Chromolaena, Condylidium, Conoclinium, Critonia, Cronquistianthus, Eutrochium, Fleischmannia, Flyriella, Hebeclinium, Koanophyllon, Mikania, and Tamaulipa.
The classification of the tribe Eupatorieae, including species placed in Eupatorium in the present or past, is an area of ongoing research, so further changes are likely. What seems fairly certain by now is that there is a monophyletic group containing Eupatorium (about 42 species of white flowered plants in North America, Europe and Asia, but not South America) and the Joe-pye weeds (Eutrochium), and possibly others.[2]
Uses
Eupatorium species are used as food plants by the caterpillars of some Lepidoptera:[verification needed]
- Bucculatrix eupatoriella (a leaf-miner) - only known from Common Boneset (E. perfoliatum)
- V-pug (Chloroclystis v-ata)
- Wormwood Pug (Eupithecia absinthiata)
- Schinia trifascia
- Coleophora case-bearers:
- C. follicularis
- C. trochilella
- C. troglodytella
A few species or cultivars, such as E. sordidum and E. coelestinum cv. 'Album', are sometimes used as ornamental plants. In particular, they are good for structural or background plantings or to attract butterflies; especially E. itatiayense[verification needed], E. maximiliani[verification needed] and E. squalidum[verification needed] are known as good honey plants.
Tobacco leaf curl virus is a pathogen occasionally affecting plants of this genus.
Medical use
Boneset, although poisonous to humans and grazing livestock, has been used in folk medicine[3], for instance to excrete excess uric acid which causes gout. Eupatorium has many more presumed beneficial uses, including treatment of dengue fever, arthritis, certain infectious diseases, migraine, intestinal worms, malaria, and diarrhea. Boneset infusions are also considered an excellent remedy for influenza. Scientific research of these applications is rudimentary at present, however.
Caution is advised when using boneset, since it contains toxic compounds that can cause liver damage. Side effects include muscular tremors, weakness, and constipation; overdoses may be deadly.
Compounds that occur in this genus include Herniarin (7-O-methylumbelliferone, 7-methoxycoumarin; in aya-pana, E. ayapana).
Selected species
North America
- Eupatorium album – White Thoroughwort[4][5]
- Eupatorium altissimum[4][5]
- Eupatorium capillifolium – "dog-fennel"[4][5]
- Eupatorium compositifolium[4][5]
- Eupatorium hyssopifolium – Hyssop-leaved Thoroughwort[4][5]
- Eupatorium lancifolium[4][5]
- Eupatorium leptophyllum[4][5]
- Eupatorium leucolepis[4][5]
- Eupatorium linearifolium (Eupatorium cuneifolium)[4][5]
- Eupatorium mikanioides[4][5]
- Eupatorium mohrii[4][5]
- Eupatorium perfoliatum L. – Common Boneset[4][5]
- Eupatorium resinosum – Pine Barren Boneset[4][5][6]
- Eupatorium rotundifolium – Round-leaved Thoroughwort[4][5]
- Eupatorium semiserratum[4][5]
- Eupatorium serotinum – Late Boneset, Late Thoroughwort[4][5]
- Eupatorium sessilifolium L. – Upland Boneset[4][5]
Europe
- Eupatorium cannabinum – Hemp-agrimony[4][5]
Asia
- Eupatorium amabile[4][7]
- Eupatorium benguetense C. Robinson[4]
- Eupatorium camiguinense Merr.[4]
- Eupatorium chinense L.[4][7]
- Eupatorium formosanum Hayata[4][7]
- Eupatorium fortunei Turcz. – fujibakama, pei lan[4]
- Eupatorium japonicum[8]
- Eupatorium lindleyanum DC.[4][7]
- Eupatorium luchuense Nakai[4][7]
- Eupatorium nodiflorum DC.[4]
- Eupatorium quaternum DC.[4]
- Eupatorium sambucifolium Elmer[4]
- Eupatorium shimadai Kitam.[4]
- Eupatorium squamosum D. Don[4]
- Eupatorium tashiroi Hayata[4][7]
- Eupatorium toppingianum Elmer[4]
- Eupatorium variabile Makino[4]
- Eupatorium yakushimaense Masam. & Kitam[4]
Uncategorized
- Eupatorium adamantiumGardner
- Eupatorium bracteatum Gardn.
- Eupatorium foeniculum
- Eupatorium itatiayense[verification needed] Hieron.
- Eupatorium laevis[verification needed] DC.
- Eupatorium makinoi[verification needed]
- Eupatorium maximiliani[verification needed] Schrad. ex DC.
- Eupatorium officinalis
- Eupatorium pacificum[verification needed]
- Eupatorium pyrifolium[verification needed] DC.
- Eupatorium rufescens[verification needed] P.W.Lund. ex DC.
- Eupatorium squalidum[verification needed] DC.
- Eupatorium urticaefolium
- Eupatorium vauthierianum[verification needed] DC.
Moved to other genera
- Eupatorium amygdalinum (moved to Ayapana)
- Eupatorium ayapana – aya-pana, Water Hemp (moved to Ayapana)
- Eupatorium coelestinum – mistflower (moved to Conoclinium)
- Eupatorium collinum (moved to Chromolaena collina)[9][10]
- Eupatorium gayanum – asmachilca (moved to Aristeguietia gayana[11][12])
- Eupatorium ligustrinum (moved to Ageratina ligustrina[13])
- Eupatorium maculatum (moved to Eutrochium maculatum) Joe-Pye Weed
- Eupatorium megalophyllum (moved to Bartlettina sordida)
- Eupatorium purpureum (moved to Eutrochium purpureum)
- Eupatorium rugosum (moved to Ageratina altissima)
- Eupatorium sordidum (moved to Bartlettina sordida)[14][15]
Footnotes
- Whittemore (1987)
- Ito et al. (2000), Schmidt & Schilling (2006)
- Sharma et al. (1999)
- a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai Schmidt & Schilling (2000)
- a b c d e f g h i j k l m n o p q r "Eupatorium". Flora of North America.
- DL Byers (1998). "Effect of cross proximity on progeny fitness in a rare and a common species of Eupatorium (Asteraceae)". American Journal of Botany 85: 644. http://www.amjbot.org/cgi/content/abstract/85/5/644.
- a b c d e f "Eupatorium". Digital Flora of Taiwan.
- "Eupatorium Linn.". Dinghushan Plant Checklist.
- "Eupatorium collinum". nomen.at.
- "Eupatorium collinum". Henriette's Herbal.
- Database entry Ayapana - Ayapana triplinervis - Ayapana - Eupatorium ayapana - Ayapana - Eupatorium triplinerve
- Fine Chem Trading (ChemFinder - UK) - Supplier MS8888
- "Eupatorium ligustrinum DC.". United States Department of Agriculture Agricultural Research Service, Beltsville Area, Germplasm Resources Information Network.
- "Eupatorium sordidum Less.". USDA PLANTS.
- Webb, C.J.; Sykes, W.R.; Garnock-Jones, P.J. (First electronic edition, Landcare Research, June 2004). "B. sordida". Flora of New Zealand. Retrieved on 2008-01-28.
References
- Ito, Motomi; Watanabe, Kuniaki; Kita, Yoko; Kawahara, Takayuki; Crawford, D.J. & Yahara, Tetsukazu (2000): Phylogeny and Phytogeography of Eupatorium (Eupatorieae, Asteraceae): Insights from Sequence Data of the nrDNA ITS Regions and cpDNA RFLP. Journal of Plant Research 113(1): 79-89. doi:10.1007/PL00013913 (HTML abstract)
- Lamont, E.E. (1995): Taxonomy of Eupatorium Section Verticillata (Asteraceae). New York Botanical Garden Press. ISBN 0-89327-391-0
- Schmidt, Gregory J. & Schilling, Edward E. (2000): Phylogeny and biogeography of Eupatorium (Asteraceae: Eupatorieae) based on nuclear ITS sequence data. Am. J. Bot. 87(5): 716-726. PMID 10811796 PDF fulltext
- Sharma, Om P.; Dawra, Rajinder K.; Kurade, Nitin P. & Sharma. Pritam D. (1999): A review of the toxicosis and biological properties of the genus Eupatorium. Natural Toxins 6(1): 1–14. doi:10.1002/(SICI)1522-7189(199802)6:1%3C1::AID-NT3%3E3.0.CO;2-E (HTML abstract)
- Whittemore, Alan (1987): The Sectional Nomenclature of Eupatorium (Asteraceae). Taxon 36(3): 618-620. doi:10.2307/1221856
Source: adapted by the editor from Wikipedia, the free encyclopedia; from the article "Eupatorium". Image Credit.